Distinguished for his researches on animal genetics, especially the study of breeding systems in relation to genetic gains in economic characters. His earlier work was on selective mating in Drosophila. Later he developed an interesting biochemical model of heterosis and studied the relationship between coincidence and crossing over. His most recent studies are on genetic control of developmental canalization in the case of the scute gene in Drosophila. His work is a major contribution to our understanding of the theoretical basis of artificial selection. In Sydney he has developed a very active research unit in animal genetics.

Expertise type

  • Biology
  • Genetics

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