Noted for the diversity and the range of his contributions to knowledge in the broad field of agricultural science with special reference to biochemistry and the animal nutrition. His earlier investigations contributed significantly to the recognition of colbalt as an essential nutrient for the sheep and of the nutritional conditions leading to infection with Clostridium botulinum in ruminants.
More recently, his ruminal flora studies in the sheep have thrown light on the effects of dietary levels of protein and starch on the nature and activity of the flora and of the amino-acid content of rumen bacterial protein. Has played a notable part in elucidating the problem of reproductive disfunction in ewes grazing subterranean clover and in demonstrating the role of clover oestrogens in this.